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and any lionesses undergo spontaneous abortions.
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This all might seem very cruel
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but from a genetic standpoint it makes sense.
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The other cubs are genetic competition for him.
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They do not have his genes.
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Moreover, only once they're out of the way
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can he reproduce and copulate with the females.
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The females do their spontaneous abortions
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because that's a reliable adaptive trick.
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These cubs are not going to survive once they are born
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so the female's best strategy
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is to get rid of them and start anew.